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This document contains excerpts taken from Chapter 9 of the book "Principles of Zoology 18th edition". It contains key points and summary which highlights the important terms and topics related to the Architectural pattern of animals.
Typology: Schemes and Mind Maps
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Zoologists today recognize 32 phyla of multicellular animals, each phylum characterized by a distinctive body plan and biological properties that set it apart from all other phyla. All are survivors of perhaps 100 phyla that appeared 600 million years ago during the Cambrian explosion, the most important evolutionary event in the geological history of life. The basic uniformity of biological organization derives from the common ancestry of animals and from their cellular construction.
The unicellular groups- are the simplest eukaryotic organisms and represent the protoplasmic grade of organization. They are nonetheless complete organisms that perform all functions of life seen in more complex animals. The multicellular animals - evolved greater structural complexity by combining cells into larger units. An animal cell is a specialized part of the whole organism and, unlike a unicellular organism, it is not capable of independent existence. The simplest metazoans show the cellular grade of organization, in which cells demonstrate division of labor but are not strongly associated with performing a specific collective function. In the more complex cell-tissue grade of organization, cells are grouped together and perform their common functions as a highly coordinated unit called a tissue. Animals at or beyond the cell-tissue grade of the organization are termed eumetazoans. In animals of the tissue-organ grade organization, tissues are assembled into still larger functional units called organs. The chief functional cells of an organ are called parenchyma. The supportive tissues are its stroma. Most animals have an additional level of complexity in which different organs operate together as organ systems.
Animal Symmetry Symmetry - refers to balanced proportions, or correspondence in size and shape of parts on opposite sides of a median plane. Spherical symmetry means that any plane passing through the center divides a body into equivalent, or mirrored, halves. This type of symmetry occurs among some unicellular forms and is rare in animals. Radial symmetry applies to forms that can be divided into similar halves by more than two plane passing through the longitudinal axis, as might be achieved when slicing a pie. These are tubular, vase, or bowl shapes found in some sponges and in hydras, jellyfish, sea urchins, and related 1
groups, in which one end of the longitudinal axis is usually the mouth. A variant form is biradial symmetry in which, because of some part that is single or paired rather than radial, only two planes passing through the longitudinal axis produce mirrored halves. Bilateral symmetry applies to animals that can be divided along a sagittal plane into two mirrored portions—right and left halves. Bilateral animals form a monophyletic group of phyla called the Bilateria. Bilateral symmetry is strongly associated with cephalization, differentiation of a head. Some convenient terms used for locating regions of bilaterally symmetrical animals are: anterior, used to designate the head end; posterior , the opposite or tail end; dorsal , the back or upper side; and ventra l, the front or belly side. Medial refers to the midline of the body; lateral , to the sides. Distal parts are farther from the middle of the body; proximal parts are nearer.
A body cavity is an internal space. The most obvious example is a gut cavity or digestive tract. Sponges , at the cellular grade of organization, do not have a gut cavity. Sponges, like all animals, develop from a zygote to a blastula stage. In animals other than sponges, development proceeds from a blastula to a gastrula stage. The depression becomes a gut cavity, also called a gastrocoel or archenteron. The external opening to the depression is the blastopore; it typically becomes the adult mouth or anus. The gut lining is endoderm and the outer layer of cells, surrounding the blastocoel, is ectoderm. 2
Segmentation , also called metamerism , is another common feature of animals. Segmentation is a serial repetition of similar body segments along the longitudinal axis of the body. Each segment is called a metamere , or somite. In forms such as earthworm s and other annelids.
Animal bodies consist of cellular components, derived from the three embryonic germ layers— ectoderm, mesoderm, and endoderm —as well as extracellular components.
In all eumetazoans, body fluids are subdivided into two fluid “compartments”: those that occupy intracellular space, within the body’s cells, and those that occupy extracellular space, outside the cells. In animals with closed vascular systems (such as segmented worms and vertebrates), the extracellular fluids are subdivided further into blood, and interstitial fluid. Interstitial fluid, also called tissue fluid, occupies the space surrounding cells (intercellular space). Many invertebrates have open circulatory systems, however, with no true separation of blood plasma from interstitial fluid.
Tissues - A tissue is a group of similar cells (together with associated cell products) specialized for performance of a common function. The study of tissues is called histology.
An epithelium (pl., epithelia) is a sheet of cells that covers an external or internal surface. Outside the body, epithelium forms a protective covering. Inside, epithelium lines all organs of the body cavity, as well as ducts or passageways through which various materials and secretions move. Simple squamous epithelium- is composed of flattened cells that form a continuous lining of blood capillaries, lungs, and other surfaces where it permits diffusion of gases and transport of other molecules into and out of cavities. Simple columnar epithelium- resembles cuboidal epithelium, but the 4
cells are taller and usually have elongate nuclei. Stratified squamous epithelium - consists of two to many layers of cells adapted to withstand mild mechanical abrasion and distortion. Junctional complexes called desmosome. Transitional epithelium- is a type of stratified epithelium specialized to accommodate great stretching. This
Connective tissues - are a diverse group of tissues that serve various binding and supportive functions. They are so widespread in the body that removal of other tissues would still leave the complete form of the body clearly apparent. Loose connective tissue , also called areolar connective tissue , is the “packing material” of the body that anchors blood vessels, nerves, and body organs. It contains fibroblasts that synthesize the fibers and ground substance of connective tissue and wandering macrophages that 5
cell or fiber , specialized for contraction. striated muscle - appears transversely striped (striated), with alternating dark and light bands. In vertebrates we recognize two types of striated muscle: skeletal and cardiac muscle. Skeletal muscle - occurs in both invertebrates and vertebrates. It is composed of extremely long, cylindrical fibers, which are multinucleate cells that may reach from one end of the whole muscle to the other. Skeletal muscle is called voluntary muscle (in vertebrates) because it contracts when stimulated by nerves under conscious central nervous system control. Cardiac muscle - is found only in the vertebrate heart. The cells are much shorter than those of skeletal muscle and have only one nucleus per cell (uninucleate). Cardiac muscle tissue is a branching network of fibers with individual cells interconnected by junctional complexes called intercalated discs. Cardiac muscle is considered involuntary muscle because it does not respond to nerves under conscious control of the central nervous system. In invertebrates , a third striated muscle type called obliquely striated muscle has been described. It has diagonally arranged muscle fibers, and studies suggest a slower contraction rate compared with other striated muscle. Smooth (or visceral) muscle - which lacks the characteristic alternating bands of the striated type, occurs in both invertebrates and vertebrates. Smooth muscle cells are long and tapering, each containing a single, central nucleus. Smooth muscle is the most common type of muscle in invertebrates , in which it serves as body-wall musculature and surrounds ducts and sphincters. cytoplasm of muscle fibers is called sarcoplasm , and contratile proteins within the fiber are myofibrils. 7
Nervous tissue - is specialized for reception of stimuli and conduction of impulses from one region to another. Two basic types of cells in nervous tissue are neurons, the basic functional unit of nervous systems, and neuroglia , a nonneuronal cell type that insulates neuron membranes and serves various supportive functions.
The most complex grades of animal organization permit, and to some extent even promote, evolution of large body size. Large size confers several important physical and ecological consequences for an organism. large animal has less surface area relative to its volume than does a small animal of the same shape. Larger size buffers an animal against environmental fluctuations; it provides greater protection against predation and enhances offensive 8