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Botany Lecture Midterm Exam Coverage, Lecture notes of Botany and Agronomy

Introduction Types of Cells Plant Tissues Plant Growth Plant Reproduction

Typology: Lecture notes

2019/2020

Available from 10/15/2024

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Botany Lecture Midterms
One of the major symptoms of plant blindness is the
tendency to overlook plants, either because of a lack of
knowledge about these organisms, their visual
homogeneity, their generally non-threatening nature or
the lack of visual cues such as movement of rapid
changes.
Application of Botany in Veterinary Medicine
- Study on phytochemicals
- Plant derived products for animal health
- Forage crops
- Ethnoveterinary therapeutic practices
- Poisonous plants
- And many more
Introduction to Botany
Levels of Biological Organization
- Life emerges through organization of various
levels
- With addition of each new level, novel
properties emerge called emergent properties
Cells as the basic unit of life
- Two distinct groups of cells exist
- Prokaryotic
- Simple and small
- Ex. bacteria
- Eukaryotic
- Possess organelles separated
by membranes
- Ex. plants, animals, fungi
Characteristics of Life
1. Order
2. regulation
3. Growth and development
4. Energy processing
5. Response to the environment
6. Reproduction
7. Evolutionary adaptation
Diversity of Life
Eukaryota
- Plantae, protista, fungi, animalia
Prokaryota
- Eubacteria, archaebacteria
Taxonomic classification
- Domain
- Kingdom
- Phylum
- Class
- Order
- Family
- Genus
- Species
Evolution of Life
1. Population with varied inherited traits
2. Elimination of individuals with certain traits
3. Reproduction of survivors
Botany as a Science
- From the greek word botane which means
plant or herb
- Scientific study of plants
Basic Concepts
- Plant metabolism is based on the principles of
chemistry and physics
- Plants must have a means of a storing and
using information
- Plants reproduce, passing their genes and
information on their descendants.
- Genes, and the information they contain,
change.
- Plants must survive in their own environment
- Plants are highly integrated organisms
- An individual plant is the temporary results of
the interaction of genes and environment
- Pants do not have decision-making capacity
Branches of Botany
Plant anatomy - the structure of plants
Plant genetics - plant heredity
Ecology - the distribution of plants throughout
the world, and why certain species grow in
certain places
Palaeobotany - plant evolution and fossil
remains
Plant physiology - plant metabolism
Plant pathology - plant diseases
Plants can be studied at many levels
Molecular level - biochemical, molecular, and
genetic functions of plants
Cellular, tissue, and organelle - anatomy and
physiology
Community and population level - interactions
with species, with other species, and with the
environment
Importance of Plants
- Photosynthesis sustains life on earth
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One of the major symptoms of plant blindness is the tendency to overlook plants, either because of a lack of knowledge about these organisms, their visual homogeneity, their generally non-threatening nature or the lack of visual cues such as movement of rapid changes. Application of Botany in Veterinary Medicine

  • Study on phytochemicals
  • Plant derived products for animal health
  • Forage crops
  • Ethnoveterinary therapeutic practices
  • Poisonous plants
  • And many more Introduction to Botany Levels of Biological Organization
  • Life emerges through organization of various levels
  • With addition of each new level, novel properties emerge called emergent properties Cells as the basic unit of life
  • Two distinct groups of cells exist
  • Prokaryotic
  • Simple and small
  • Ex. bacteria
  • Eukaryotic
  • Possess organelles separated by membranes
  • Ex. plants, animals, fungi Characteristics of Life
  1. Order
  2. regulation
  3. Growth and development
  4. Energy processing
  5. Response to the environment
  6. Reproduction
  7. Evolutionary adaptation Diversity of Life Eukaryota
  • Plantae, protista, fungi, animalia Prokaryota
  • Eubacteria, archaebacteria Taxonomic classification
  • Domain
  • Kingdom
  • Phylum
  • Class
  • Order
  • Family
  • Genus
  • Species Evolution of Life
  1. Population with varied inherited traits
  2. Elimination of individuals with certain traits
  3. Reproduction of survivors Botany as a Science
  • From the greek word botane which means plant or herb
  • Scientific study of plants Basic Concepts
  • Plant metabolism is based on the principles of chemistry and physics
  • Plants must have a means of a storing and using information
  • Plants reproduce, passing their genes and information on their descendants.
  • Genes, and the information they contain, change.
  • Plants must survive in their own environment
  • Plants are highly integrated organisms
  • An individual plant is the temporary results of the interaction of genes and environment
  • Pants do not have decision-making capacity Branches of Botany ● Plant anatomy - the structure of plants ● Plant genetics - plant heredity ● Ecology - the distribution of plants throughout the world, and why certain species grow in certain places ● Palaeobotany - plant evolution and fossil remains ● Plant physiology - plant metabolism ● Plant pathology - plant diseases Plants can be studied at many levels ● Molecular level - biochemical, molecular, and genetic functions of plants ● Cellular, tissue, and organelle - anatomy and physiology ● Community and population level - interactions with species, with other species, and with the environment Importance of Plants
  • Photosynthesis sustains life on earth
  • Fundamental source of food
  • Medicines
  • Provide fuel, shelter, and paper products
  • Conservation biology is a critical area of research
  • Biotechnology seeks to develop new plant products Eukaryotic cells
  • Cell is the basic unit of life
  • All living organisms are made up of one or more cells
  • All cells are produced from pre-existing cells
  • Prokaryotes vs eukaryotes
  • All cells share four common components
  • Plasma membrane
  • Cytoplasm
  • Genetic material
  • Ribosomes CHARACT ERISTICS

EUKARYOTIC

CELL

PROKARYOTIC

CELL

Definition Any cell that contains a clearly defined nucleus and membrane bound organelles Any unicellular organism that does not contain a membrane bound nucleus or organelles Examples Animal, plant, fungi, and protist cells Bacteria and archaea Nucleus Present (membrane bound) Absent (nucleoid region Cell size Large (10-100μ) Small (less than a micrometer to 5 μm) DNA replication Highly regulated with selective origins and sequences Replicates entire genome at once Organism type Usually multicellular Unicellular Chromoso mes More than one One long single loop of DNA and plasmids Ribosomes Large Small Growth rate/ generation time Slower Faster Organelles Present Absent Ability to store hereditary information All eukaryotes have this ability All prokaryotes have this ability Cell wall Simple, present in plants and fungi Complex, present in all prokaryotes Plasma membrane Present Present Cytoplasm present Present CELLULA R PROCESS

ORGANELL

E

FUNCTION

Manufactur ing nucleus Controls cellular processes; store genetic material ribosomes Site of protein synthesis endoplasmic reticulum (smooth ER or rough ER) Assembles lipids (SER) and proteins (RER)

  • Responsible for manufacturing and storing of food Chloroplast
  • Surrounded by two membranes
  • Outer membrane is permeable to small organic molecules; inner membrane is less permeable and studded with transport proteins
  • The innermost matrix of chloroplasts, called the stroma contains metabolic enzymes and multiple copies of the chloroplast genome
  • Third internal membrane called the thylakoid membrane, which is extensively folded and appears as stacks of flattened disks in electron micrographs
  • Contain the light harvesting complex including pigments such as chlorophyll, as well as the electron transport chains used in photosynthesis Central vacuole
  • An organelle that occupies 30-90% of the cell’s volume
  • Used to adjust the cell’s size and turgor pressure
  • May contain digestive enzymes
  • May serve as storage compartments for nutrients and metabolites Plasmodesmata
  • Singular = plasmodesmata
  • Small channels that directly connect the cytoplasm of neighboring plant cells to each other, establishing living bridges between cells Cell cycle
  • Living organisms reproduce by two methods
  • Asexual
  • Offspring are identical to the original cell or organism
  • Involves inheritance of all genes from one parent
  • Sexual
  • Offspring are similar to parents but show variations in traits
  • Involves inheritance of unique sets of genes from two parents
  • Series of events from the time a cell forms until its cytoplasm divides
  • Includes three phases: interphase, nuclear division (e.g. mitosis), cytoplasmic division Cytokinesis differs for plant and animal cells
  • Cytokinesis
  • Cleavage in animal cells
  • Cleavage furrow forms from a contracting ring of microfilaments, interacting with myosin
  • Cleavage furrow deepens to separate the contents into two cells
  • Cytokinesis in plant cells
  • Cell plate forms in the middle from vesicles containing cell wall material
  • Cell plate grows outward to reach the edges, dividing the contents into two cells
  • Each cell has a plasma membrane and cell wall

Meristems and Plant Tissues Meristem

  • the plant tissue composed of totipotent cells that allows plant growth
  • Centers of mitotic cell division, and are composed of a group of undifferentiated self-renewing stem cells from which most plant structure arise
  • Responsible for keeping the plant growing Classified by their location ● Apical - root and shoot tips ● Lateral - vascular and cork cambia ● Intercalary - at internodes, stem regions between the places at which leaves attach, and leaf bases, especially of certain monocotyledons Meristematic Tissues ● Apical
  • Found at, near, or the tips of roots and shoots
  • Increase plant in length (primary growth)
  • 3 primary meristems: protoderm, ground meristem, procambium ● Lateral
  • Increase the growth of roots and stems (secondary growth)
  • Vascular cambium (for support and conduction)
  • Cork cambium (in the form of a thin cylinder that runs the length of roots and stems of woody plants ● Intercalary
  • Develop at intervals along stems, where, like the tissues produced by apical meristems, their tissues add to stem length Primary meristem
  • Responsible for primary growth (via apical meristem)
  • Three primary meristems
  • Protoderm ~ epidermis
  • Ground meristem ~ core (ground)
  • Phloem
    • Specialized to move organic materials from sources to sinks ● Ground tissue
  • Bulk of the plant body
  • Food production, storage, support
  • Lies between dermal and vascular tissue
  • Eudicot stem ground tissue is divided into pith and cortex
  • Leaf ground tissue is called mesophyll Primary Growth: Shoot and Root Systems Plants as a vascular organism
  • Nonvascular plants
  • No vessels
  • No stems
  • No leaves
  • No roots
  • Vascular plants
  • Have vessels for transport of water (xylem) and food (phloem)
  • Have roots, stems, and leaves
  • Absorbs water and minerals from soil through roots
  • Absorb the sun’s energy and carbon dioxide from the air through shoots (stems and leaves)
  • Plant roots depend on shoots for carbohydrates produced via photosynthesis
  • Plant shoots depends on roots for water and mineral
  • Autotroph ~ self feeding Roots *when a seed germinates, a part of the embryo within it (radicle), grows out and develops into the first root
  • Functions
  • Anchorage
  • Storage
  • Absorption
  • Conduction

Types of root systems

  • Taproot
    • Develop from an enlarged primary root
    • Functions
      • Storage of food
      • Regulate the formation and growth of branch roots
      • Reach deep underground water sources
  • Fibrous
    • Form from masses of adventitious roots that replace the primary root
    • Special adaptations: may hold soil particles *most dicot plants have taproot systems *monocots have fibrous root systems Root structure
  • Root regions
  • Internal morphology Four regions or zones in developing young root
  • Region of maturation
  • Region of elongation
  • Region of cell division
  • Root cap Root cap
  • Composed of a thimble shaped mass of parenchyma cells covering the tip of each root
  • Has no equivalent in stems
  • Functions
  • Protection from damage of delicate tissues
  • Perception against gravity (amyloplast) Region of cell division
  • Composed of an apical meristem
  • Cells divide every 12 to 36 hours
  • Cells in this region is mostly cube shaped
  • The apical meristem soon subdivides into three meristematic areas: protoderm, ground tissue, procambium Region of elongation
  • Usually extends about 1cm or less from the tip of the root
  • No further increase in cell size takes place above this region Region of maturation
  • Root hairs adhere tightly to soil particles
  • Root hairs are not separate cells
  • Cortex ~ food storage
  • Root cortex is similar to that of the stem except for the presence of endodermis
  • Primary cell walls of the cells in the endodermis are impregnated with suberin
  • Suberin forms bands (casparian strip) around the walls perpendicular to the root’s surface
  • Having the central stem of the plant to grow dominantly over the other parts/sides of the stem
  • Control by the shoot tip over axillary buds Leaf structure
  • Lamina ~ leaf blade
  • Petiole ~ attached the leaf to the stem
  • Midrib ~ associated with the vasculature
  • Venation ~ arrangement of the veins Leaf structure and function
  • Epidermis ~ for the regulation of gas exchange
  • Upper and lower epidermis
  • Contains the stomata, each stoma surrounded by two guard cells
  • With waxy cuticle to reduce water loss
  • May have other adaptations such as having trichomes
  • Mesophyll
  • Inner tissue (parenchyma) of a leaf, containing many chloroplasts
  • Palisade parenchyma/mesophyll ~ column shaped, tightly packed
  • Spongy parenchyma/mesophyll ~ loosely arranged irregularly-shaped cells Leaf Characteristics
  • Phyllotaxy (alternate, spiral, opposite)
  • Leaf form (simple, compound) Leaf venation
  • Parallel
  • Netted
  • Dichotomous (ginkgo) *plants exhibit indeterminate growth where the plant will continue adding new organs (leaves, stems, roots) as long as it has access to necessary resources. This is because they have meristematic tissues Secondary Growth Secondary growth from lateral meristems
  • Increases the girth of organs
  • Provides protection for expanding organs
  • Develops a supporting system to allow for arborescent growth Vascular cambium (lateral meristem)
  • Accounts for most secondary growth
  • Lied between the xylem and phloem in stems and roots
  • Forms from the procambium in vascular tissue
  • Arises from parenchyma between the vascular bundles to form a complete ring of vascular cambium
  • Consists of a thin layer of parenchyma cells Secondary Xylem
  • Forms the wood of the stem
  • Wood can be classified as
  • Spring wood
  • Thin walled and large diameter elements
  • Summer wood
  • Thicker walled and smaller diameter
  • Differences in spring and summer wood form distinct growth rings
  • Usually one per year so called annual rings
  • Can provide information about the life cycle of the tree
  • Relative wetness or dryness of each season
  • Age of the tree
  • Secondary growth can continue over thousands of years Wood Softwood
  • Composed of tracheids, no fibers
  • Has rays composed of sheets of cells one cell by 20 cells high
  • Resin canals often in rays Hardwood
  • Tracheids, vessel, many fibers
  • Fibers make wood hard
  • Ring-porous wood has large vessels only in early wood
  • Diffuse-porous wood has large vessel throughout the wood
  • Ray structure is diverse, but usually rays are 30 cells wide and hundreds of cells high *ray ~ a band of usually parenchymatous cells extending from the cambium into both the xylem and phloem of a plant root or stem that conducts fluids radially and appears in a cross section like a spoke of a wheel Sapwood
  • Outer few centimeters
  • Water and solutes move in it
  • Light, pale, and relatively structurally weak Heartwood
  • Center wood of the stem
  • Wastes are removed here
  • Fills with tannins, resins, gums, and oils
  • Darker, denser, and more aromatic than sapwood Secondary Phloem
  • Formed from fusiform initials in the vascular cambium towards the outer surface of the tree
  • Transports water and dissolved organic materials
  • Contains sieve elements and parenchyma alternating with bands of thick walled fibers
  • Innermost sieve elements functional

Structure of Flowers

  • Each flower begins as an embryonic primordium that develops into a bud, and occurs as a specialized branch at the tip of a stalk (peduncle). The peduncle swells at the tip into a small pad (receptacle)
  • Flowers typically contain four types of highly modified leaves called floral organs - Sepals ~ enclose and protect flower bud - Petals ~ showy; attract pollinators - Stamen ~ male reproductive structures - Pistil ~ female reproductive structures *complete flower ~ if a flower has stamen, pistil and sepal *perfect flower ~ if flower contains both functional stamen and pistils Stamen
  • Anther ~ produces pollen, which house cells which develop into sperm
  • Filament ~ elevates anther Carpel
  • Stigma ~ site of pollination
  • Style ~ “neck” that leads to ovary
  • Ovary ~ houses ovules, which contain developing egg Carpel
  • The female reproductive organ of a flower, consisting of an ovary, a stigma, and usually a style
  • May occur singly or as one of a group Pistil
  • Composed of an ovary, one or more styles (which may be absent), and one or more stigmas

Life Cycle of a generalized angiosperm Plant Life Cycle: Alternating Generations

  • Involves alternating diploid (2n) and haploid (n) generations - The diploid generation ~ sporophyte - Specialized diploid cells in anthers and ovules undergo meiosis to produce haploid spores - Haploid spores undergo mitosis and produce the haploid generation - The haploid generation is called the gametophyte Male Gametophyte: Pollen Grain - A cell in the anther undergoes meiosis to produce four haploid spored - Each spore divides via mitosis to produce two cells called the tube cell and generative cell - A tough wall forms around the cells to produce a pollen grain - Pollen grains are released from the anther Female Gametophyte: Embryo Sac - A cell in the ovule undergoes meiosis to produce four haploid spores - Three of the spores degenerate - The surviving spore undergoes a series of mitotic divisions to produce the embryo sac - One cell within the embryo sac is an egg ready for fertilization - One central cell within the embryo sac has two nuclei and will produce endosperm Germination and Fertilization - Pollen grain germination - Tube nucleus produces pollen tube, which grows down through the style to the ovary - Generative nucleus divides to produce two sperms - Double fertilization - One sperm fertilizes the egg to produce a zygote - One sperm fuses with the central cell nuclei to produce 3n endosperm
  • Radicle ~ embryonic root *plumule and radicle are enclosed in a tubular, sheathing structure called coleoptile and coleorhiza, respectively Germination
  • Interplay of many factors
  • Many require dormancy
  • Seed coats
  • One way valve
  • Inhibitors for survival Seed germination continues life cycle
  • Germination breaks seed dormancy
  • Germination begins when water is taken up