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Botany Lecture Final Exam Coverage, Lecture notes of Botany and Agronomy

Plant Hormones Water Transport Phylogeny and Systematics

Typology: Lecture notes

2019/2020

Available from 10/15/2024

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Botany Lecture Finals
Plant Hormones
Five major types of hormones regulate plant
growth and development
- Hormone is chemical signal that is
produced in one part of the body and
transported to another, where it
triggers responses in target cells
- Binding of hormones to specific
cellular receptors triggers a signal
transduction pathway
- Tiny amounts of hormone can have a
big effect
Auxin stimulates the elongation of cells in
young shoots
- Indoleacetic acid (IAA) ~ naturally
occurring auxin that promotes
seeding elongation
- Produced in shoot apical meristems
and transported downward through a
plant
- Concentration of auxin and site of
activity are important to auxin’s
effects
- In moderate concentration, auxin
promotes cell elongation in stems
- In high concentrations, auxin reduces
cell elongation in stems
- Affects cell elongation in roots at
lower concentration
Cytokinins stimulate cell division
- Promote cytokinesis or cell division
- Are produced in actively growing
organs such as roots, embryos, and
fruits
- Produced in roots then move upward
through the plant
- Retard aging in leaves and flowers
Gibberellins affect stem elongation and have
numerous other effects
- Promote stem elongation by
increasing cell division and
elongation
- Also promote fruit development and
seed germination
- Act antagonistically against another
plant hormone called abscisic acid
Abscisic acid inhibits many plant processes
- Inhibits growth
- High concentration ~ promote seed
dormancy
- ABA must be removed for
germination to occur
- The ratio of ABA to gibberellins
controls germination
- Influences plant water relations
- Accumulation of ABA in
wilted leaves promotes
stomatal closure
- ABA produced in roots can
signal low soil moisture
conditions and triggers plans
to conserve water by closing
stomata
Ethylene triggers fruit ripening and other
aging processes
- Is a gaseous by product of natural
gas combustion and a naturally
occurring plant hormone
- Plants produce ethylene in response
to stresses such as mechanical
pressure, injury, infection, and
drought or flood
Responses to stimuli
Tropisms orient plant growth toward or
away from environmental stimuli
Tropism ~ responses that causes plants to
grow in response to environmental stimuli
- Positive tropism cause plants to grow
toward a stimulus
- Negative tropism cause plants to
grow away from a stimulus
Plants respond to various environmental
stimuli
- Phototropism ~ response to light
- Gravitropism ~ response to gravity
- Thigmotropism ~ response to touch
Plants have internal clocks
-Circadian rhythms are innate
biological cycles of approx. 24 hours
- Both plants and animals have
circadian rhythms
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Plant Hormones Five major types of hormones regulate plant growth and development

  • Hormone is chemical signal that is produced in one part of the body and transported to another, where it triggers responses in target cells
  • Binding of hormones to specific cellular receptors triggers a signal transduction pathway
  • Tiny amounts of hormone can have a big effect Auxin stimulates the elongation of cells in young shoots
  • Indoleacetic acid (IAA) ~ naturally occurring auxin that promotes seeding elongation
  • Produced in shoot apical meristems and transported downward through a plant
  • Concentration of auxin and site of activity are important to auxin’s effects
  • In moderate concentration, auxin promotes cell elongation in stems
  • In high concentrations, auxin reduces cell elongation in stems
  • Affects cell elongation in roots at lower concentration Cytokinins stimulate cell division
  • Promote cytokinesis or cell division
  • Are produced in actively growing organs such as roots, embryos, and fruits
  • Produced in roots then move upward through the plant
  • Retard aging in leaves and flowers Gibberellins affect stem elongation and have numerous other effects
  • Promote stem elongation by increasing cell division and elongation
  • Also promote fruit development and seed germination
  • Act antagonistically against another plant hormone called abscisic acid Abscisic acid inhibits many plant processes
  • Inhibits growth
  • High concentration ~ promote seed dormancy
  • ABA must be removed for germination to occur
  • The ratio of ABA to gibberellins controls germination
  • Influences plant water relations
  • Accumulation of ABA in wilted leaves promotes stomatal closure
  • ABA produced in roots can signal low soil moisture conditions and triggers plans to conserve water by closing stomata Ethylene triggers fruit ripening and other aging processes
  • Is a gaseous by product of natural gas combustion and a naturally occurring plant hormone
  • Plants produce ethylene in response to stresses such as mechanical pressure, injury, infection, and drought or flood Responses to stimuli Tropisms orient plant growth toward or away from environmental stimuli Tropism ~ responses that causes plants to grow in response to environmental stimuli
  • Positive tropism cause plants to grow toward a stimulus
  • Negative tropism cause plants to grow away from a stimulus Plants respond to various environmental stimuli
  • Phototropism ~ response to light
  • Gravitropism ~ response to gravity
  • Thigmotropism ~ response to touch Plants have internal clocks
  • Circadian rhythms are innate biological cycles of approx. 24 hours
  • Both plants and animals have circadian rhythms
  • Circadian rhythms are influenced by environmental cues such as light, but they are controlled by biological clocks
  • The biological clocks of plants are likely the result of rhythmic production of proteins that influence gene expression Plants mark the seasons by measuring photoperiod
  • Flowering, seed germination, and dormancy are all seasonal phenomena in plants
  • Detect season by measuring photoperiod, the relative lengths of day and night
  • Plant flowering signals are determined by night length
  • Short day plants flower when the dark period is greater than some critical length
  • Long day plants flower when dark period is shorter than some critical length
  • Experiments that altered light and dark periods were used to determine that it is night length and not day length that cues plants to flower Phytochrome is a light detector that may help set the biological clock
  • Phys constitute a large receptor family sensitive to red and far red light occur implants, (cyano)bacteria and fungi
  • Pr is naturally produced during dark hours, while Pfr is broken down
  • The relative amounts of Pr and Pfr present in a plant change as day length changes Water Transport in Plants Molecular movement Molecules and ions are constantly random motion
  • Diffusion (passive)
  • Solutes from higher concentration to lower concentration
  • Maintain equilibrium
  • Osmosis (passive)
  • Solvent movement from higher to lower
  • Active transport
  • Solutes from low to high concentration
  • Assisted by proteins/pumps that require energy (ATP) to open
  • Turgor
  • Have enough water inside the central vacuole
  • Maintain the size and shape of the cell
  • Plasmolysis
  • Loss of water through osmosis, accompanied by the shrinkage of protoplasm away from the cell wall
  • Imbibition
  • The stomata regulate transpiration and gas exchange
  • Changes in turgor pressures in the guard cells, which contain chloroplasts, occur when they are exposed to changes in light intensity, carbon dioxide concentration, or water concentrations
  • Stomata are open during the day and closed at night, but water is conserved in a number of desert plants whose stomata open only at night when there is less water stress on plants (CAM photosynthesis)
  • High humidity reduces transpiration and low humidity accelerates it
  • If a cool night follows a warm humid day, water droplets may be produced through hydathodes at the tips of veins of the leaf ~ guttation Plant Metabolism Autotrophs are the producers of the biosphere
  • Able to make own food without organic molecules derived from any other living thing
  • Photoautotroph ~light
  • Lithoautotroph
  • Chemoautotroph Photosynthesis
  • Takes place in chloroplasts and other parts of green organisms
  • Light energy is stored in simple sugar molecule
  • Chloroplasts
  • consisting of photosynthetic pigments, enzymes, and other molecules grouped together in membranes
  • Concentrated in the cells of the mesophyll, the green tissue in the interior of the leaf
  • Two membranes encloses the stroma, the dense fluid within the chloroplast
  • A system of interconnected membranous sacs called thylakoids segregates the stroma from another compartment, the thylakoid space
  • Thylakoids are concentrated in stacks called grana
  • Chlorophyll resides in the thylakoid membranes The principal components of photosynthesis Water
  • Less than 1% is absorbed and used
  • Source of oxygen
  • Limiting factor in photosynthesis Light
  • Exhibits properties of both waves and particles
  • About 40% of radiant energy we receive is in the form of visible light Chlorophyll
  • Most plants have two kinds
  • 𝛼 ~ bluish green C35H72MgN4O
  • 𝛽 ~ yellowish green C55H70MgN4O
  • 𝛼 is three times more than 𝛽 usually
  • Other pigments
  • Carotenoids ~ yellow to orange
  • Phycobillins ~ blue or red
  • Chlorophyll a, d, e Photosynthesis is a redox process Two major steps in photosynthesis
  • Light reaction and calvin cycle Light reaction
  • Light energy converted in the thylakoid membranes to chemical energy and O
  • Water is split to provide O2 as well as electrons
  • H+ ions reduce NADP+ to NADPH

Each pigment has its own distinctive pattern of light absorption Pigment absorption spectrum

  1. Pigment absorbs light
  2. Energy levels of some of the pigments electrons are raised
  3. Emitted also in light Each pigment has its own distinctive pattern of light absorption
  • Pigments, molecules that absorb light are built into the thylakoid membrane
  • Chloroplasts contain several different pigments and all absorb light of different wavelengths
  • Chlorophyll a absorbs blue violet and red light and reflects green
  • Chlorophyll b absorbs blue and orange and reflects yellow-green
  • The carotenoids absorb mainly blue-green light and reflect yellow and orange Photosystems
  • The energy released could be lost as heat or light, but rather it is conserved as it is passed from one molecule to another molecule Photosystems capture solar power
  • All of the components to accomplish this are organized in thylakoid membranes in clusters called photosystems
  • Photosystems are light harvesting complexes surrounding a reaction center complex
  • Pigment absorbs photons and pass the energy to the reaction center
  • Each photosynthetic unit of Photosystem I consists of chlorophyll a and b, carotenoid and a pair of special reaction-center molecule of chlorophyll P
  • Each photosynthetic unit of Photosystem II consists of chlorophyll a and b, B-carotene and reaction-center molecule of chlorophyll P Overview of the two stages of photosynthesis in a chloroplast Electron flow in the light reactions
  1. A pigment molecule in a light-harvesting complex absorbs a photon of light. The energy is passed on until it reaches the reaction center of Photosystem II to excite electron of chlorophyll P
  2. The electron is captured in the primary electron acceptor
  3. Water is split, its electron electrons are supplied one by one to P680,
  • A cladogram depicts patterns of shared characteristics among taxa
  • A clade is a group of species that includes an ancestral species and all its descendants
  • Cladistics studies resemblances among clades
  • A shared derived character (apomorphies) is an evolutionary novelty unique to a particular clade Importance of naming plants and classifying them
  • To impart information about the plant
  • Necessary for communication
  • Common names
  • Native names
  • International names Division Bryophyta: Mosses
  • Gametophyte typically consists of small leaflike structures (not true leaves) arranged spirally or alternately around a stem-like axis
  • The axis is anchored to its substrate by means of rhizoids
  • Leaves are the only one cell thick (except at the midrib), lack vascular strands and stomata and are haploid Division of Anthocerophyta: Hornworts
  • Found in tropical forests, along streamsides, and in disturbed fields around the world
  • The group’s common name “hornwort” refers to the tall narrow sporophytes which are embedded in the top of the plant (gametophyte) Division Hepaticophyta: Liverwort
  • “Wyrt” means plant
  • “Liver plant” because flattened gametophyte resembles the liver
  • Stomata fixed open
  • Gametophyte stage is dominant stage in both liverworts and hornworts
  • Viewed as the plants most closely related to the ancestor that moved to land Division Lycopodiophyta: Club mosses, Spike mosses, Quillworts
  • Plants with true leaves, roots, and stems
  • Leaves are, without exception, microphylls (one single, unbranched leaf vein) and this is a defining feature of the group
  • Roots are adventitious

Division Psilophyta: Whisk ferns

  • The only living vascular plants to lack both roots and leaves
  • Stem lacks roots, it is anchored instead by a horizontally creeping stem called a rhizome
  • The erect position of the stem bears paired enations, outgrowths which looks like miniature leaves but unlike true leaves, the enations have no vascular tissue Division Equisetophyta: Horsetail
  • With a single extant family
  • Whorled microphylls and hollow, jointed, green stems
  • Some species have numerous small branches and bear a slight resemblance to a horse’s tail Division Polypodiophyta: Ferns
  • A group of non seed plants
  • With true leaves, roots, and stems Gymnosperms and Angiosperms Gymnosperms Cone-bearing
  • “naked seeds”
  • Separate male and female gametes
  • Pollination by wind
  • With vascular system
  • Seeds not enclosed in ovary but enclosed in cones or modified leaves
  • Classified into four divisions
  • Coniferophyta (same pattern of seed development and with secondary cambium)
  • Cycadophyta (same pattern of seed development and with secondary cambium)
  • Ginkgophyta(same pattern of seed development and with secondary cambium)
  • Gnetophyta (produced true xylem tissue) Gymnosperm Life Cycle
  • Involves alternation of generation with a dominant sporophyte in which the female gametophyte resides
  • Male and female reproductive organs can form cones
  • Male cones (staminate) give rise to pollen grains by meiosis
  • Pollen grain houses two sperm cells
  • Yellow pollen is released and carried by the wind during spring
  • Some gametophytes will land on female cone
  • One sperm will unite with egg
  • Upon fertilization, diploid egg will give rise to embryo Coniferophyta/Pinophyta
  • Dominant group of gymnosperms
  • Most variety of species
  • Typically tall trees that usually bear scale-like or needle-like leaves (adaptation)
  • Predominant at high altitudes and cold climates
  • Many coniferous trees are harvested for paper, pulp and timber
  • “softwood” – contains tracheids but no vessel elements Cycadophyta
  • Thrive in mild climates
  • Often mistaken for palms
  • Large cones commonly pollinated by beetles rather than wind (unusual for gymnosperm)
  • Some face possible extinction
  • Often used as ornamental plants
  • Rarely branched trunks with soft pithy wood Ginkgophyta
  • Fan-shaped leaves
  • Male and female organs are produced on separate plants
  • Deciduous
  • Leaves alternate and simple
  • “Living fossil” with no close relatives
  • Single surviving species: Maidenhair tree G. biloba Gnetophyta
  • Closest relative to modern angiosperm
  • Includes three orders with one genera each (~70 sp.): Ephedra, Gnetum and Welwitschia
  • Broad leaves
  • Vines or small shrubs in subtropical regions
  • Posses vessel elements in their xylem